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Pseudomonas species, a large class of gram-negative bacteria, exhibit significant biomedical, ecological, and industrial importance. Pseudomonas, a major human pathogen, is able to cause severe infections in vulnerable patients hospitalized with cystic fibrosis, burns, acquired immunodeficiency syndrome, or cancer. Extensive research has been focused to dissect the molecular basis of the infection mechanisms and to develop novel therapeutic means against Pseudomonas infections. The genetic manipulation in Pseudomonas is extremely urgent.